In addition to processing mail via SMTP and the Store driver (Think Get/Push mail from mailbox databases) Exchange will process mail messages placed in a folder on the hard disk of a HubTransport server. Known as the "pickup folder" it is not configured by default in Exchange 2010 sp1.
Set-TransportServer -identity ExchHub01 -pickupfolder "C:\incomingMail\"
For more information on configuration of the pick up folder see:
http://technet.microsoft.com/en-us/library/bb124549.aspx
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